aDepartment of Biology, Faculty of Mathematics and Natural Sciences, University of Brawijaya, Jl Veteran, Malang, East Java, Indonesia bDepartment of Biotechnology and Medical Engineering, Faculty of Biosciences and Medical Engineering, Universiti Teknologi Malaysia, Johor Bahru, Malaysia cDepartment of Biotechnology, Faculty of Science, IIUM, Bandar Indera Mahkota, Kuantan, Pahang, Malaysia